Biomedical Sciences (BMS) Graduate Program
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Dr. Didier Stainier
BMS Research Areas
BMS Faculty Directory 
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All of the faculty associated with the BMS graduate program are listed below. Click on a faculty name to view a detailed description of an individual research program, contact information, recent publications and links to other relevant websites. Each of our faculty is associated with one or more of our eight thematic areas; click on the thematic area name listed by each of our faculty to read more about the research going on in that area, and to view a list of all faculty associated with that thematic area.

A
Abul Abbas, MBBS
Primary Research Interest: Immunology
T-cell regulation in tolerance and immunity

Nina Agabian, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Immunology
Microbial Pathogenesis

Nadav Ahituv, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Tissue/Organ Biology and Endrocrinology
Gene regulation and human disease

Tamara Alliston , PhD
Primary Research Interest: Tissue/Organ Biology and Endrocrinology
Secondary Research Interest: Developmental and Stem Cell Biology
Skeletal Cell Differentiation and Matrix Quality

Arturo Alvarez-Buylla, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Developmental & Stem Cell Biology
Birth and Migration of New Neurons

Mark Anderson, MD, PhD
Primary Research Interest: Immunology
The Genetic Control of Autoimmune Disease

Raul Andino, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Replication of RNA Viruses and Vaccine Development

K. Mark Ansel, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Developmental and Stem Cell Biology

Regulation of gene expression in the immune system

Kaveh Ashrafi, PhD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Neurobiology & Human Genetics
Genetics  and Neurobiology of Fat and Feeding Regulation

B
Steinunn Baekkeskov, PhD
Primary Research Interest: Immunology
Molecular Mechanisms of Autoimmunity/Structure, Cell Biology, and Immunopathology of Autoantigens

Anna Bakardjiev, MD
Primary Research Interest: Virology & Microbial Pathogenesis
Host-Pathogen Interactions at the Maternal-Fetal Interface

Allan Balmain, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Mouse models for the study of human cancer susceptibility and cancer progression

Scott Baraban, PhD
Primary Research Interest: Neurobiology
Neurobiology of Epilepsy

Diane Barber, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Signal Transduction and Cytoskeleton Organization

Jody Baron, MD, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
Innate and adaptive immunity to viral pathogens

Allan Basbaum, PhD
Primary Research Interest: Neurobiology
The Neurological Basis of Pain and Its Control

Michael S. Beattie, PhD
Primary Research Interest: Neurobiology
CNS injury and repair

Hilary Beggs, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Neurobiology
Cell-Matrix-Cytoskeletal Signaling: Mechanisms of Eye Development and Disease

Gabriele Bergers, PhD
Primary Research Interest: Vascular & Cardiac Biology
Secondary Research Interest: Cancer Biology & Cell Signaling / Developmental & Stem Cell Biology
Interactions between tumor cells and blood vessels to regulate neovascularization, tumor growth and invasion

Harold Bernstein, MD, PhD
Primary Research Interest: Vascular & Cardiac Biology
Secondary Research Interest: Developmental & Stem Cell Biology
Cell Cycle Regulation in Cardiovascular Biology

Daniel Bikle, MD, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Regulation of cellular differentiation and function by calciotropic hormones

J. Michael Bishop, MD
Primary Research Interest: Cancer Biology & Cell Signaling
Molecular Genetics of Normal and Neoplastic Cells

Montgomery Bissell, MD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Cancer Biology & Cell Signaling
Liver Wound Repair

Brian Black, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Vascular & Cardiac Biology
Transcriptional control of mammalian organogenesis

Elizabeth Blackburn, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Cancer Biology & Cell Signaling
Telomeres and Telomerase

Robert Blelloch, MD, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Cancer Biology & Cell Signaling
Molecular basis of stem cell self-renewal, differentiation and contribution to cancer

Jeffrey Bluestone, PhD
Primary Research Interest: Immunology
Regulation of T-cell activation and tolerance during autoimmune and transplantation responses

Nancy Boudreau, PhD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Cancer Biology & Cell Signaling
Role of Homeobox genes in tumor induced angiogenesis and wound healing

V. Courtney Broaddus, MD
Primary Research Interest: Cancer Biology & Cell Signaling
Apoptotic Synergy in Mesothelioma

Frances Brodsky, DPhil
Primary Research Interest: Immunology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Intracellular Membrane Traffic: Mechanisms & Immune Function

Katja Brückner, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Cancer Biology & Cell Signaling
Drosophila hematopoiesis and epithelial plasticity- models for cancer development

Benoit Bruneau, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Vascular & Cardiac Biology
Transcriptional regulation of heart development

Laura Bull, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Genetics approaches to study of the biology of the liver in health and disease

Nigel Bunnett, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Protease switches for inflammation and pain signals

Esteban Gonzalez Burchard, MD, MPH
Primary Research Interest: Human Genetics
Identifying ethnic specific genetic and biologic risk factors for asthma and asthma severity among U.S. ethnic minority groups

C
Pat Calarco, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Molecular and Cellular Analysis of Preimplantation Mammalian Development

George Caughey, MD
Primary Research Interest: Immunology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Genetics, Structure and Function of Mast Cell Secretory Proteases

Harold Chapman, MD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Integrin signaling in lung fibrosis and cancer

Israel F. Charo, MD, PhD
Primary Research Interest: Immunology
Structure and Function of Chemokine Receptors

Ben Cheyette, MD, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Developmental & Stem Cell Biology
Wnt Signaling in Development and Function of the Mammalian Brain

Pao-Tien Chuang, MD, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Hedgehog Signaling in Mammalian Embryogenesis and Postnatal Physiology

James Cleaver, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Human Genetics
DNA Repair Mechanisms and Human Disease

Philip Coffino, MD, PhD
Primary Research Interest: Human Genetics
Proteasomes, substrate recognition and processing

Fred Cohen, MD, DPhil
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Neurobiology
Protein Folding and Drug Design

Bruce Conklin, MD
Primary Research Interest: Vascular & Cardiac Biology
Secondary Research Interest: Developmental & Stem Cell Biology
Rewiring Receptor and G-protein Signals In Vivo

David Copenhagen, PhD
Primary Research Interest: Neurobiology
Synapses, Circuits and Development in Retina

Joe Costello, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Human Genetics
Integrated genomic and epigenomic mechanisms contributing to cancer

Shaun Coughlin, MD, PhD
Primary Research Interest: Vascular & Cardiac Biology
Secondary Research Interest: Cancer Biology & Cell Signaling
Signaling mechanisms in vascular biology and disease

Lisa Coussens, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Immunology
Manipulating the Host Response to Cancer

Jeffery Cox, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Immunology
Host-Pathogen Interactions Responsible for M. tuberculosis Virulence

Lindsey A. Criswell, MD, MPH
Primary Research Interest: Human Genetics
Secondary Research Interest: Immunology
Genetics and epidemiology of human autoimmune disease

Jason Cyster, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
Molecular Regulation of Cell Migration and Survival in Lymphoid Tissues

D
Mary Dallman, PhD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Brain-Pituitary-Adrenal Interrelationships

Creg Darby, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Human Genetics
Biofilms of pathogenic bacteria; C. elegans models of bacterial pathogenesis

Jayanta Debnath, MD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
The Role and Regulation of Autophagy in Epithelial Homeostasis and Cancer

Anthony DeFranco, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Regulation of B Lymphocyte and Macrophage Function by Cell Surface Receptors

Joseph DeRisi, PhD
Primary Research Interest: Human Genetics
Genomic approaches to yeast molecular biology and infectious disease

Rik Derynck, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Cancer Biology & Cell Signaling
Transmembrane TGF-a and TGF-b Receptor Signaling in Cell Proliferation and Differentiation

Ivan Diamond, MD, PhD
Primary Research Interest: Neurobiology
Adenosine and the Neurobiology of Alcoholism

Marc Diamond, MD
Primary Research Interest: Neurobiology
Secondary Research Interest: Human Genetics
Molecular Mechanisms of Neurodegeneration in Polyglutamine Expansion Diseases

E
Robert Edwards, MD
Primary Research Interest: Neurobiology
Molecular Analysis of Neurotransmitter Release

Joanne Engel, MD, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Immunology
Interaction of Bacterial Pathogens with their Host Eukaryotic Cells

David Erle, MD
Primary Research Interest: Immunology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Immunopathogenesis of asthma; immune cell trafficking

Gerard Evan, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Human Genetics
Molecular Basis of Carcinogenesis: Regulation of Cell Proliferation and Cell Death (Apoptosis)

F
Robert V. Farese, Jr., MD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Vascular & Cardiac Biology
Cell Biology of Energy Metabolism

Donna Ferriero, MD
Primary Research Interest: Neurobiology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Injury Response in the Developing Nervous System

John Fike, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Radiation, neurogenesis and cognitive function

Steven Finkbeiner, MD, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Cancer Biology & Cell Signaling
Molecular Mechanisms of Synaptic Plasticity and Neurodegeneration

Susan Fisher, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Vascular & Cardiac Biology
Placental Biology/Bacterial Adhesion

Lawrence Fong, MD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Immunobiology of Dendritic Cells in Disease

Alan Frankel, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
RNA-Protein Recognition

Ying-Hui Fu, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Neurobiology
Human genetics in neurodegenerative diseases

G
Don Ganem, MD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Cancer Biology & Cell Signaling
Biology and Replication of Human Viral Pathogens

Fen-Biao Gao, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Neurobiology
Neuronal Development and Neurodegeneration

David Gardner, MD
Primary Research Interest: Vascular & Cardiac Biology
Growth control in the cardiovascular system: Role of the natriuretic peptides and their receptors

Adam Gazzaley, MD, PhD
Primary Research Interest:
Neurobiology
Neural Mechanisms of Attention and Memory

Michael German, MD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
The Biology of Pancreatic Beta Cells and Diabetes

Kathleen Giacomini, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Cancer Biology & Cell Signaling
Pharmacogenomics of Membrane Transporters

Jane Gitschier, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Neurobiology
Genetic Defects in Heavy Metal Transport

Linda C. Giudice, MD, PhD, MSc
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Normal and abnormal endometrial development

Ed Goetzl, MD
Primary Research Interest: Immunology
Immunoregulatory G Protein-Coupled Receptors of T-Cells

Andrei Goga, MD, PhD
Primary Research Interest: Cancer Biology and Cell Signaling
Secondary Research Interest: Developmental and Stem Cell Biology, Human Genetics
Oncogenes and Cell Cycle Control

Douglas B. Gould, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Vascular & Cardiac Biology
Genetic and molecular mechanisms of ocular and cerebrovascular diseases

Warner Greene, MD, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Immunology
Molecular Analysis of HIV Pathogenesis

Carl Grunfeld, MD, PhD
Primary Research Interest: Vascular & Cardiac Biology
Secondary Research Interest: Immunology
Metabolic Changes and the Host Response to Infection

H
Steven Hamilton, MD, PhD
Primary Research Interest: Human Genetics
Genetics and Pharmacogenetics of Psychiatric Disorders

Douglas Hanahan, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Human Genetics
Modelling Pathways to Cancer: Cellular, Genetic & Immunological Determinants, and Experimental Therapeutics

Hobart Harris, MD, MPH
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
Lipoproteins and Immune Responses to Infection and Acute Injury

Ulrike Heberlein, PhD
Primary Research Interest: Neurobiology
Molecular genetics of addiction-related behaviors in flies and mice

Matthias Hebrok, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Cancer Biology & Cell Signaling
Molecular mechanisms of pancreas development, function and disease

Graeme Hodgson, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Cancer Biology & Cell Signaling
Gene Discovery and Targeted Molecular Therapeutics in Brain Tumors

Wen-Chi Hsueh, MPH, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Genetic epidemiology of aging- and metabolism- related complex traits in humans

Eric Huang, MD, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Developmental & Stem Cell Biology
Molecular Mechanisms Controlling Sensory Neuron Development

Yadong Huang, MD, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Vascular & Cardiac Biology
Molecular Mechanisms and Therapeutic Strategies of Neurodegenerative Diseases

I
Holly Ingraham, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Cancer Biology & Cell Signaling
Mechanisms of Hormone Action in Organ Development and Disease


J
Robert Jaffe, MD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Growth and Differentiated Function in Human Fetal Adrenal and Gonad

C. David James, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Human Genetics
CNS Tumor Molecular Biology and Therapeutic Testing

Lily Jan, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Vascular & Cardiac Biology
Potassium Channels

Alexander Johnson, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Human Genetics
Virulence and Evolution of a Human Pathogen

David Julius, PhD
Primary Research Interest: Neurobiology
Molecular Biology of Neurotransmitter Receptors and Ion Channels

K
Nigel Killeen, D.Phil
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
The determination of T-cell fate

Ophir Klein, MD, PhD

Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Human Genetics
Developmental genetics of dental and craniofacial structures

Jane Koehler, MD
Primary Research Interest: Virology & Microbial Pathogenesis
Pathogenic Interactions Between the Hemotrophic Bacterium Bartonella and Mammalian Host Cells

Scott Kogan, MD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Developmental & Stem Cell Biology
Cooperating Events in Leukemogenesis and Their Reversal

Tom Kornberg, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Human Genetics
Organogenesis and patterning during development

Anatol Kreitzer, PhD
Primary Research Interest: Neurobiology
Synaptic plasticity and circuit function in the basal ganglia

Arnold Kriegstein, MD, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Neurobiology
Neural Stem Cells and Embryonic Cortical Development

Matthew Krummel, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Molecular Coordination in Multipartite Interactions

Pui-Yan Kwok, MD, PhD
Primary Research Interest: Human Genetics
Analysis of Complex Genetic Traits

L
Diana Laird, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Human Genetics, Tissue/Organ Biology & Endocrinology
Molecular and cellular mechanisms of primordial germ cell development

Lewis Lanier, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Regulation of T cell and NK cell immune responses by activating and inhibitory receptors

Corey Largman, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Developmental & Stem Cell Biology
Mechanism of HOX Homeodomain Protein Regulation of Cell Growth and Transformation

Jennifer LaVail, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Virology & Microbial Pathogenesis
Axonal Transport Mechanisms of Herpers Simplex Virus in CNS Neurons

Andrew Leavitt, MD
Primary Research Interest: Developmental & Stem Cell Biology
Megakaryocytopoiesis and platelet function

Jay Levy, MD
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
HIV Pathogenesis

Joachim Li, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Regulation of Eukaryotic DNA Replication

Limin Liu, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Immunology
Metabolism and Regulation of Nitric Oxide Bioactivity

Richard Locksley, MD
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
Development of effector T lymphocytes

Stavros Lomvardas, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Developmental & Stem Cell Biology
Identification and characterization of stochastic choices in the nervous system

Clifford Lowell, MD, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Tyrosine Kinase Regulated Signal Transduction in Blood Cells

M
Averil Ma, MD, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Innate Immunity and Immunological Memory

Tippi C. MacKenzie, MD
Primary Research Interest: Immunology
Secondary Research Interest: Developmental & Stem Cell Biology
Tolerance induction after in utero stem cell transplantation or gene therapy

Hiten Madhani, MD, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Human Genetics
How a good yeast lives and how a bad yeast kills:  S. cerevisiae vs. C. neoformans

Jacquelyn Maher, MD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Vascular & Cardiac Biology
Tissue-protective cytokines; fatty liver disease

Marta Margeta, MD, PhD
Primary Research Interest: Neurobiology
Molecular determinants of neuronal susceptibility to degeneration

Todd Margolis, MD, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Neurobiology
Ocular Infection with the Herpes Virus

Gail Martin, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Vertebrate Organogenesis

Mehrdad Matloubian, MD, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
Lymphocyte Migration and Autoimmune Disease

Theodora Mauro, MD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Developmental & Stem Cell Biology
Ionic Fluxes Control Keratinocyte Adhesion, Differentiation and Secretion

Frank McCormick, PhD, FRS
Primary Research Interest: Cancer Biology & Cell Signaling
Therapeutic Approaches to Cancer

Joseph McCune, MD, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Immunology
Pathogenic Mechanisms, Treatment, and Prevention of HIV Disease

Steven McIntire, MD, PhD
Primary Research Interest: Neurobiology
Genetic Analysis of Synaptic Function in C. Elegans

James McKerrow, MD, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Immunology
Biochemistry, Immunopathogenesis and Molecular Biology of Parasitic Diseases

Martin McMahon, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Oncogenes, Signal Transduction and Cancer

Michael McManus, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Developmental & Stem Cell Biology , Immunology
Mammalian microRNA and RNA Interference Pathways

Synthia Mellon, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Neurobiology
Developmental Regulation, Molecular Biology and Novel Actions of Steroid/Neurosteroid Synthesis

Takashi Mikawa, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Vascular & Cardiac Biology
Developmental Regulation of Organogenesis

Keith Mostov, MD, PhD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Developmental & Stem Cell Biology
Morphogenesis and function of epithelial organs

Paul Muchowski, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Human Genetics
Molecular mechanisms that underlie neurodegenerative disorders

Lennart Mucke, MD
Primary Research Interest: Neurobiology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Molecular Neurobiology of CNS Diseases

John Murnane, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Mechanisms of genomic instability in mammalian cells

N
Laura Napolitano, MD
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
The Regulation of T Lymphopoiesis

Stephen Nishimura, MD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Cancer Biology & Cell Signaling
Mechanistic basis of integrin function homeostasis

Robert Nissenson, PhD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Cancer Biology & Cell Signaling
Signal Transduction Mechanisms in Bone Cell Differentiation

Douglas Nixon, MD, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
Cellular immune responses to HIV

Linda Noble, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest:
Neurobiology of CNS Injury

Robert Nussbaum, MD
Primary Research Interest: Human Genetics
Secondary Research Interest: Neurobiology
Human Genetic Analysis of Parkinson's Disease and Other Disorders

O
Scott Oakes, MD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Immunology
Control of Intrinsic Pathways of Apoptosis by the BCL-2 Family of Proteins

Jeffrey Olgin, MD
Primary Research Interest: Vascular & Cardiac Biology
Mechanisms of cardiac arrhythmias

Charles Ordahl, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Embryonic Muscle Development

Melanie Ott, MD, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Immunology

Molecular Pathogenesis of HIV and Hepatitis C Virus Infections

P
Joel Palefsky, MD
Primary Research Interest: Virology & Microbial Pathogenesis
DNA tumor viruses of the epithelium

Feroz Papa, MD, PhD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Developmental & Stem Cell Biology
Protein misfolding diseases and novel therapies

Emmanuelle Passegué, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Immunology
Molecular and cellular understanding of normal and leukemogenic hematopoiesis

David Pearce, MD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Vascular & Cardiac Biology
Regulation of gene expression and epithelial ion transport by stress steroids.

B. Matija Peterlin, MD
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
Molecular biology of MHC II and HIV

Samuel Pleasure, MD, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Developmental & Stem Cell Biology
Control of early hippocampal development

Louis Ptacek, MD
Primary Research Interest: Human Genetics
Secondary Research Interest: Neurobiology
Genetics and biology of human nervous system disorders

Jennifer Puck, MD
Primary Research Interest:
Immunology
Secondary Research Interest: Human Genetics
Inherited disorders of lymphocyte development


R
Miguel Ramalho-Santos, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Regulation of Stem Cell Pluripotency and Reprogramming

Louis Reichardt, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Developmental & Stem Cell Biology
Extracellular Factors Affecting Neuron Development


Jeremy Reiter, MD, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Cancer Biology and Cell Signaling, Human Genetics, Vascular & Cardiac Biology
Signaling at the Primary Cilium, the Cell's Antenna

Neil Risch, PhD

Primary Research Interest: Human Genetics
Human Genetics, Population Genetics, Statistical Genetics, Genetic Epidemiology


Dorit Ron, PhD

Primary Research Interest: Neurobiology
Signal transduction and neurobiology of addiction

Jeroen Roose, PhD
Primary Research Interest: Cancer Biology and Cell Signaling
Secondary Research Interest: Immunology
Ras Signal Transduction in Lymphocytes and Cancer

Steve Rosen, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Roles of biological sulfation in leukocyte trafficking and tumorigenesis

Philip Rosenthal, MD

Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Human Genetics
Molecular, translational, and clinical studies of malaria

David H. Rowitch, MD, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Neurobiology
CNS Development and Disease


John Rubenstein, MD, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Developmental & Stem Cell Biology
Genetic Regulation of Regional Specification and Differentiation in the Forebrain

Davide Ruggero, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Human Genetics
Dynamics of translation control in gene expression, cancer, and disease


Eric Rulifson, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Neurobiology
Developmental Genetics of the Drosophila Neuroendocrine System and Diabetes

S
Rich Schneider, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Craniofacial Morphogenesis and Evolution

Bill Seaman, MD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
The Role of Lymphocyte Subtypes in Immunity
 

Neil Shah, MD, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Developmental & Stem Cell Biology
Targeted Therapy of Hematologic Malignancies

Nirao Shah, MD, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Human Genetics, Developmental & Stem Cell Biology
Neural control of sexually dimorphic behaviors

Kevin Shannon, MD

Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Human Genetics
Molecular Genetics and Therapy of Myeloid Leukemia


Robin Shaw, MD, PhD
Primary Research Interest: Vascular & Cardiac Biology
Secondary Research Interest: Cancer Biology & Cell Signaling
Regulation of Cardiac Ion Channels: Cytoskeleton and Membrane Targeting


Dean Sheppard, MD

Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Vascular & Cardiac Biology, Tissue/Organ Biology & Endocrinology
In vivo function of integrins and molecular basis of lung diseases

Jeoung-Sook Shin, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Virology & Microbial Pathogenesis
Antigen presentation by dendritic cells and its role in the pathogenesis of asthma

Anita Sil, MD, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Human Genetics
Fungal pathogenesis and host response


David Sretavan, MD, PhD

Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Neurobiology
Mechanisms of axon injury, disease, and novel methods of axon repair

Deepak Srivastava, MD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Human Genetics, Vascular & Cardiac Biology
Molecular and genetic dissection of cardiogenesis and its role in human disease

Didier Stainier, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Vascular & Cardiac Biology
Zebrafish Developmental Genetics/Cardiovascular Development

Paul M. Sullam, MD
Primary Research Interest: Virology & Microbial Pathogenesis
Pathogenetic mechanisms of Gram-positive pathogens

Raymond Swanson, MD
Primary Research Interest: Neurobiology
Astrocyte-neuron Interactions in Nervous System Disease

T
Qizhi Tang, PhD
Primary Research Interest: Immunology
Regulatory T cell control of autoimmunity and transplant rejection

Laurence Tecott, MD, PhD
Primary Research Interest: Neurobiology
Molecular Genetic Analysis of Serotonin Systems and Complex Behavioral Traits

Thea Tlsty, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Regulation of Genomic Instability in Mammalian Cells

David Toczyski, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Activation of the DNA damage checkpoint

U
Erik Ullian, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Developmental & Stem Cell Biology
The Role of Glia in the Formation and Function of Synaptic Circuits

V
Christian Vaisse, MD, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Genetics of Metabolic Diseases

Eric Verdin, MD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Immunology
HIV Pathogenesis and lymphocyte biology

Zena Vexler, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Vascular & Cardiac Biology, Immunology
Inflammatory mechanisms of injury and repair after adult and neonatal stroke

Susan Voglmaier, MD, PhD
Primary Research Interest: Neurobiology
Vesicle Trafficking and Neurotransmitter Release

Thiennu Vu, MD, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
The regulation of lung vascularization and development

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Matthias Wabl, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Somatic Cell Genetics of the B Lymphocyte

Rong Wang, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Vascular & Cardiac Biology
Signaling in Embryonic and Tumor Angiogenesis

Valerie Weaver, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Developmental & Stem Cell Biology
The tissue microenvironment and mechanical force in adult and stem cell fate

Orion Weiner, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Vascular & Cardiac Biology
Cell polarity during chemotaxis

Richard Weiner, PhD
Primary Research Interest: Neurobiology
Secondary Research Interest: Cancer Biology & Cell Signaling
Neuroendocrine Control of Reproduction and Signaling Pathways for the Antiangiogenic Factor 16K PRL

Art Weiss, MD, PhD
Primary Research Interest:
Immunology
Secondary Research Interest: Cancer Biology & Cell Signaling
Signal Transduction Events and Lymphocyte Activation

William Weiss, MD, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Secondary Research Interest: Human Genetics
Mouse Models for Tumors of the Nervous System

Zena Werb, PhD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Cancer Biology & Cell Signaling
Functions of the Extracellular Microenvironment in Development and Cancer

Joe Wiemels, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Cancer Biology & Cell Signaling
Etiology and genetics of childhood leukemia and brain cancer

Holger Willenbring, MD
Primary Research Interest: Developmental & Stem Cell Biology
Secondary Research Interest: Cancer Biology & Cell Signaling

Mechanisms of liver regeneration and cancer

John Witte, PhD
Primary Research Interest: Human Genetics
Genetic Epidemiology of Cancer and other Complex Diseases

Torsten Wittmann, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Cytoskeletal Dynamics and Cell Polarization

David Wofsy, MD
Primary Research Interest: Immunology
Mechanisms of autoimmunity

Paul Wolters, MD
Primary Research Interest: Immunology
Secondary Research Interest: Tissue/Organ Biology & Endocrinology
Cellular and molecular pathogenesis of lung disease

Anthony Wynshaw-Boris, MD, PhD
Primary Research Interest: Human Genetics
Secondary Research Interest: Developmental & Stem Cell Biology
, Neurobiology
Pathogenesis of Human Neurogenetic Diseases

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Allison Xu, PhD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Neurobiology
Hypothalamic regulation of energy balance, obesity and type 2 diabetes

Y
Keith Yamamoto, PhD
Primary Research Interest: Cancer Biology & Cell Signaling
Signal Transduction and Transcriptional Regulation

Hal Yee, Jr, MD, PhD
Primary Research Interest: Tissue/Organ Biology & Endocrinology
Secondary Research Interest: Cancer Biology & Cell Signaling
The Biology of Tissue Pericytes/Myofibroblasts and their Role in Fibrotic Diseases

Ben Yen, MD, PhD
Primary Research Interest: Virology & Microbial Pathogenesis
Secondary Research Interest: Cancer Biology & Cell Signaling
Regulation of Gene Expression by Hepatitis B Virus

Midori Yenari, MD
Primary Research Interest: Neurobiology
Secondary Research Interest: Vascular & Cardiac Biology, Immunology
Inflammatory mechanisms in cerebral ischemia

Z
Scott Zamvil, MD, PhD
Primary Research Interest: Immunology
Secondary Research Interest: Neurobiology, Human Genetics
Pathogenesis and immune modulation of CNS autoimmunity

Elad Ziv, MD
Primary Research Interest: Human Genetics
Application of Population Genetics to the Study of Complex Traits in Humans

Dr. J. Michael Bishop, Chancellor, UCSF
Dr. Elizabeth Blackburn
Dr. Elizabeth Blackburn
Dr. Lennart Mucke
Dr. Lennart Mucke

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